Chevin Chase veterinary practice is buzzing with excitement. The finishing touches are being made to the marquee in the garden with flowers arriving and tables being laid. Eight months pregnant Anna is directing the activity but begins to panic. Time is ticking away and her father George, the groom, is still operating on a puppy. Meanwhile in London, Sarah walks out on her cheating boyfriend David and drives up north. The church is packed to the rafters. Practice Manager Deborah is making disparaging remarks about everyone to her husband Rick, the practice Senior Partner. During a brief lull in her tirade, he slips outside for a sneaky cigarette with nurse Matt. Meanwhile fellow nurses Clare and Margaret enjoy a gossip about the guests and Sarah's imminent arrival. Sarah hasn't been home since her mum died two years ago and everyone wonders why... not least her sister Anna. Little does she know that Sarah hides a painful secret - she caught Anna's husband Tom in bed with one of the nurses, Fiona. And she hasn't been able to face any of them since. Just as Sarah enters the church, Fiona takes her seat. There's no time for trouble as the young bride Claudie, 30 years George's junior, makes her breath-taking entrance. At the wedding reception, chemistry ignites between Rick and Clare who share a brief kiss. Meanwhile, George makes a shocking announcement - he's retiring and moving to Malta with wife Claudie, leaving pregnant Anna and Sarah with a struggling business, one vet down, one vet defecting to the local opposition and a list of operations as long as their arm. George's revelation is enough to send Anna into labour. Anna and Tom race to the hospital, leaving Sarah in charge of the practice and their unruly kids. Anna gives birth to a girl who she names after her mother, Imogen. But George drops another shocking bombshell - they've re-mortgaged the house by £200,000. This leaves Sarah with no option but to muck in at the Chase. After a heated discussion, Fiona and Sarah make amends. Fiona reveals she has a son Theo from a previous failed relationship but little does Sarah know that the child is Tom's. Sarah's astonished when David turns up, begging her to come home, but she reveals she is home. Elsewhere, Tom's late on Theo's monthly payment. Fiona threatens to reveal all if he doesn't cough up soon...

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The staff are stunned by the death of a loved colleague. Anna contemplates life as a single mother while Sarah must give an answer to Adrian's proposal.

The sisters are infuriated by the arrival of Claudie's sister Paula, and the contents of her beauty salon. There are other new arrivals in Oxley as both Fiona's father and Montgomery's younger brother appear to have an interest in the Williams family. But nothing can prepare them for the final revelation of the day, as the police come knocking at the door.

As the investigation into George's murder begins, Anna and Sarah are distracted by the possibility of losing everything to their stepmother.

The atmosphere at The Chase becomes strained, and police, poodles and pamper packs culminate in a shock resignation.

With morale at an all-time low, the staff look forward to attending the Montgomery Estate ball, where Anna finds an unlikely prince charming.

It's the morning after the ball and Anna is nowhere to be found. Wondering where she is, Tom takes matters into his own hands. Just as it seems they might be reconciled, Fiona strikes again. Sparks fly, and Anna's response to Tom's infidelity is finally made clear. The practice is in chaos with Margaret on reception and Paula's clients taking over, plus Claudie's brother Liam and his dog Stan. Anna finds comfort from a surprising source.

Tom is reeling from Fiona's confession and the realisation he is again embroiled in her life. Meanwhile, the local battle of the bands leaves Matt with a tricky dilemma and he and Anna are forced to consider their relationship. Sarah turns to Sebastian for support when she discovers Adrian knows the truth about her fertility problems and that Anna betrayed her trust.

There's excitement at The Chase as Tom treats a lion cub on the premises. Anna contemplates her new relationship with Matt, while Sarah agonises over her one-night stand with Sebastian. Claudie is facing sentencing for her fraud conviction and is panicked at the possibility of being sent to jail. A quick earner for Liam has dangerous and expensive ramifications.

Deborah is in her element as she moonlights as William Montgomery's PA. As their friendly rapport develops, Deborah wonders if their relationship could become something other than professional. Margaret's internet dating gets her involved with a local hunt saboteur. Rick oversteps the mark with his jealousy and lashes out at Clare. Could this finally spell the end of their affair?