Chevin Chase veterinary practice is buzzing with excitement. The finishing touches are being made to the marquee in the garden with flowers arriving and tables being laid. Eight months pregnant Anna is directing the activity but begins to panic. Time is ticking away and her father George, the groom, is still operating on a puppy. Meanwhile in London, Sarah walks out on her cheating boyfriend David and drives up north. The church is packed to the rafters. Practice Manager Deborah is making disparaging remarks about everyone to her husband Rick, the practice Senior Partner. During a brief lull in her tirade, he slips outside for a sneaky cigarette with nurse Matt. Meanwhile fellow nurses Clare and Margaret enjoy a gossip about the guests and Sarah's imminent arrival. Sarah hasn't been home since her mum died two years ago and everyone wonders why... not least her sister Anna. Little does she know that Sarah hides a painful secret - she caught Anna's husband Tom in bed with one of the nurses, Fiona. And she hasn't been able to face any of them since. Just as Sarah enters the church, Fiona takes her seat. There's no time for trouble as the young bride Claudie, 30 years George's junior, makes her breath-taking entrance. At the wedding reception, chemistry ignites between Rick and Clare who share a brief kiss. Meanwhile, George makes a shocking announcement - he's retiring and moving to Malta with wife Claudie, leaving pregnant Anna and Sarah with a struggling business, one vet down, one vet defecting to the local opposition and a list of operations as long as their arm. George's revelation is enough to send Anna into labour. Anna and Tom race to the hospital, leaving Sarah in charge of the practice and their unruly kids. Anna gives birth to a girl who she names after her mother, Imogen. But George drops another shocking bombshell - they've re-mortgaged the house by £200,000. This leaves Sarah with no option but to muck in at the Chase. After a heated discussion, Fiona and Sarah make amends. Fiona reveals she has a son Theo from a previous failed relationship but little does Sarah know that the child is Tom's. Sarah's astonished when David turns up, begging her to come home, but she reveals she is home. Elsewhere, Tom's late on Theo's monthly payment. Fiona threatens to reveal all if he doesn't cough up soon...

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The staff are stunned by the death of a loved colleague. Anna contemplates life as a single mother while Sarah must give an answer to Adrian's proposal.

The sisters are infuriated by the arrival of Claudie's sister Paula, and the contents of her beauty salon. There are other new arrivals in Oxley as both Fiona's father and Montgomery's younger brother appear to have an interest in the Williams family. But nothing can prepare them for the final revelation of the day, as the police come knocking at the door.

As the investigation into George's murder begins, Anna and Sarah are distracted by the possibility of losing everything to their stepmother.

The atmosphere at The Chase becomes strained, and police, poodles and pamper packs culminate in a shock resignation.

With morale at an all-time low, the staff look forward to attending the Montgomery Estate ball, where Anna finds an unlikely prince charming.

Deborah is back at work and enforcing a tough new regime. Faced with pedometers and mission statements, Clare and Rick are feeling the strain. Matt saves the day for Anna as he covertly stops Claudie making more changes to the house.

Tempers become heated at The Chase as Deborah fires Clare while Rick stands helplessly by. Anna and Sarah move from one crisis to the next as the mediation over George’s estate leaves Claudie holding all the cards. Tom, desperate to escape Fiona's mind games, makes a last-ditch attempt to persuade Anna to move to Botswana. Rick is devastated to see Clare with a mystery man.