It all began ten years ago. D2s attacking Boston, a blood-soaked concert hall: The incident known as the Boston Tragedy. The conductor who lost his life in that tragedy was none other than Kenji Asahina, Takt's father. After losing his father, the boy decides to devote himself to playing the piano and he lets his rage play through his melodies. The one who took care of him was a girl named Cosette, who looked just like Destiny.

After separating from Lenny and Titan, Takt and crew continue their journey under the blazing sun. During their trip, they end up fighting some D2s and end up bringing a train on the transcontinental railroad to a sudden stop. They then see a Musicart in white armor and a Musicart with a bewitching smile. The man accompanying the two Musicarts is named Shindler and calls himself the Top Conductor of the New York Symphonica.

Anna and Destiny happen to see a town destroyed during their travels. D2s are awakening everywhere and attacking various towns and people, reverting the world back to four years ago, before Sagan's Declaration. As the threat from the D2s grew, Takt was obsessing over something. With a blank staff notation in front of him, Takt plays an imaginary keyboard as he desperately tries to create something.

D2s awakening. Takt's hometown being attacked, and Cosette's death. This was all a part of Sagan's plan. Shindler wants to get rid of all those inferior to him in this desolate world. After learning the truth about Cosette's death, Takt charges at Shindler in a blind rage. But Takt is injured by Hell. As she witnesses Takt injured and bleeding, Destiny feels some emotions welling up inside of her that she's never felt before...