Will encourages the Glee Club to support Artie, and asks each of them to perform in a wheelchair. Meanwhile, Kurt and Rachel compete for a big solo.

Fresh off their monumental win at Sectionals, New Directions is on top of the world. With an extra spring in their step, they get to work on new numbers for regionals. But when Rachel and Finn’s budding romance hits a bump in the road after she becomes involved with a competitor, Jesse St. James, it threatens to derail the entire club. Mr. Schuester pays a visit to Shelby Corcoran, coach of Vocal Adrenaline, to confirm Jesse’s motives. Meanwhile, Will is newly single, but is he ready to take his relationship with Emma to the next level? And to top it all off, Sue Sylvester returns from her condo in Boca with a renewed focus on destroying Will and disbanding the glee club.

When a sore throat affects Rachel's singing voice, she goes into panic mode. Meanwhile, Kurt tries to change his persona to impress his father, and Puck makes a strategic move to elevate his social status.

Mr. Schuester encourages the glee club to express themselves using the music of Lady Gaga. Meanwhile, Rachael has a life-changing encounter, and Tina embraces her inner Goth.

When Brittany and the gang try to convince Mr. Schuester to let them do a Britney Spears number, he is reluctant to go that edgy. Meanwhile, an insecure Will is curious about Emma’s new beau, Dr. Carl Howell, so he convinces her to bring him in to talk to the kids about dental hygiene. While the dreamy doc inspires the girls to take their dental health more seriously, Will is left feeling worse than before.

After Finn and Sam find an unconventional way of controlling their urges, a few glee club members pick up on the strategy and start using it too. But when Coach Beiste gets wind of what they are up to, the glee club has some explaining to do. Meanwhile, the Girls vs. Boys Tournament returns, Kurt meets an ally from a rival glee club, Puck and Artie join forces and coach Beast is on the verge of quitting her job.

When Mr. Schuester gets sick, a substitute teacher takes over his Spanish class and the glee club and winds up winning over the whole New Directions clan. Meanwhile, when Sue fills in for Principal Figgins as McKinley’s top dog, she tries to flex her power on campus.

New Directions heads to the big Sectionals competition to go head-to-head against the Warblers and the Hipsters. While trouble brews between some of the young couples, Emma and Carl grow closer.

It's Christmas time at McKinley High and Finn tries to get everyone in the holiday spirit; Brittany needs no help because she still believes in Santa Claus. In other news, Sue scams the teacher's Secret Santa

A local football championship game takes place and the "Thriller" mash-up is performed during the halftime show. The Cheerios, McKinley High's cheerleading team, compete at Regionals.

Love is in the air at McKinley High when the glee club sets up a kissing booth for Valentine's Day. Things don't go as planned, however, and hearts end up getting broken.

Rachel tries to launch a comeback into the McKinley High social stratosphere with a little help from Brittany. Meanwhile, Sam starts a tribute band.

Principal Figgins teaches the student body about the danger of underage drinking, and the glee club is personally touched by the message. Meanwhile, Mr. Schuester and Coach Beiste indulge in a little fun of their own.

Holly Holiday returns as the sex education substitute teacher and helps the glee club out with some new numbers. On the other hand, Emma becomes the head of the celibacy club.

With Sue Sylvester coaching Aural Intensity, Blaine and the Warblers’ dynamic set list and tough judges Tammy Jean Albertson and Sister Mary Constance, the kids of New Directions decide the only way to win Regionals is to write and perform their own original songs. Meanwhile, Quinn is hell-bent on getting Finn back and ruling McKinley once again.

The glee club agrees to help raise funds for another under-the-radar group; big decisions are made and relationships are tested.

The glee club learns about self-acceptance through the music of Lady Gaga; Lauren and Quinn compete for prom queen; Ms. Pillsbury faces with her issues; Rachel makes a life-changing decision; Santana schemes.

Sue Sylvester revives the school newspaper and decides to print a few inflammatory blind items. The glee club gains a whole new respect for Sam when they discover some interesting new information about his family life. Meanwhile, April Rhodes (guest star Chenoweth) returns to Lima and tries to convince Mr. Schuester to hit the road with her.

When the entertainment for prom falls through, Principal Figgins asks New Directions to perform at the dance much to prom coordinator Sue’s dismay. Things get heated as the competition for prom king and queen draws closer, and Jessie St. James returns.

New Directions continues to strategize their set list for Nationals with a little help from Jessie St. James. Meanwhile, an unexpected loss rocks the world of one of McKinley’s own.

New Directions heads to Nationals to compete in the ultimate singing showdown.

The glee kids are down in the dumps following their disappointing showing at Nationals, so Will assigns them a project to get them back on track. Meanwhile, Ms. Sylvester wants to go to Washington: Sue's running for Congress.

Emma gets a visit from her parents; Mike Chang's parents get involved when his grades aren't up to their standards.

McKinley’s Sadie Hawkins dance offers several gleeks a rare chance to feel empowered. Sam and Blaine investigate a possible cheating scandal at Sectionals. Kurt makes a new friend at NYADA.

All of Will’s hard work and dedication to the Glee Club and to fostering the talents and dreams of his students pays off in a major way, and everyone shares in the feeling of joy that permeates the school as teachers and students reflect on their shared journey. Old rifts and wounds are healed and new sights are set on a promising future. Everyone seems to be ending up exactly as expected and right where they belong… along with more than a few unexpected surprises.