When Senator Perrin makes his stand against the Rossum Corporation, Echo attempts to stop his expose. Adelle, and Topher begin to see what they are up against in the rival of Washington DC.

Echo and Bennett have a shocking meeting as Adelle goes head-to-head with the ruthless head of the DC Dollhouse. Topher sees double when he must involve Victor in his espionage. Perrin finds a surprising witness to testify against the Rossum Corporation, but find he may be living in a house of cards.

Topher races to reconstruct Echo's original personality and memories when Adelle realizes they are the key to averting a devastating future. Meanwhile, Echo and Bennett's shared past and the identity of The Rossum Corporation's mysterious leader are revealed.

In the year 2020, Echo and her surviving Dollhouse crew attempt to restore order to a devastating future world before mankind is eliminated.