Dotty persuades Moley and Mishmosh to head out for a spot of camping, meanwhile the Gardener has his own plans.

When Mishmosh falls ill before the greatest inventor competition, Moley and Dotty compete in his place.

When Moley helps take care of the worms at Dotty's Adopt Worm Rescue Center, the Gardener sends Squirm to pose as a homeless worm and help him steal Manny.

When Moley spends the day with a young mole named Cheese who wants to be just like him, Moley helps Cheese recognize his own talents and uniqueness.

When MishMosh’s lab is flooded, Moley invites MishMosh to temporarily stay at his burrow. But when MishMosh proves to be a difficult houseguest by turning Moley’s place into a makeshift lab, Moley’s patience is tested.

Mrs. Moley is directing a small play, with Moley cast as the hero, but his fear gets the better of him.

When the Gardener finally gets his hands on Manny, Moley and Dotty are on his trail, so he dresses up as the world’s greatest Mole detective and lead them away from the clues.