On his first day, rookie officer Ben Sherman is assigned to veteran cop John Cooper, his no-nonsense training officer. After being involved in a fatal shooting, Ben already begins questioning his career choice. Detectives Sammy Bryant and Nate Moretta try to convince a young teen to testify after learning she is an eyewitness in the attempted murder of a boy who was mistaken for a gang member. Meanwhile, Detective Lydia Adams investigates the disappearance of a young girl.

An investigation is opened after Ben, John, and Officer Chickie Brown encounter an infant crawling down the street. Lydia takes a personal interest in the case after Child Protective Services gets involved, and goes out on a limb for the mother. Meanwhile, Sammy and Nate are furious when they learn their teenage eyewitness won't get police protection due to lack of funds, and put up their own money after an attempt is made on her life.

Ben and John are called to the home of a man who wants them to find his dog, and another in which a drug overdose led to homicide. Chickie and her partner, Officer Bill "Dewey" Dudek, butt heads after he makes some poor decisions that may have repercussions, and later, Dewey finds himself in an embarrassing situation.

A young Jane Doe found murdered in an alley sets Lydia on an emotional journey, and she discovers her suspect was involved in a cold case. Meanwhile, Sammy's wife, Tammi, decides to embark on a new career, and Nate deals with some family issues and reveals a deep secret.

In an effort to recover Detective Daniel "Sal" Salinger's stolen weapon, Sammy and Nate crack down on the residents in the neighborhood where it occurred. Elsewhere, a man exposes himself to Lydia.

Lydia and her partner, Detective Russell Clarke, investigate a string of jewelry thefts in Bel Air and Ben discovers one of the victims is someone he knows. In another matter, Ben must face his past when it comes back to haunt him. Meanwhile, John reveals a secret he's been harboring for a long time, Russell has troubles at home, and Sammy and Nate take extraordinary steps to protect their eyewitness, Janilla, after another attempt is made on her life.

When Sammy discovers Marta has made bail, and fearing another attempt will be made on Janilla's life, he arranges to have her stay with Lydia, but Janilla's safety is compromised after she visits her old neighborhood. Meanwhile, Sal continues to spy on his daughter's activities, Dewey flips out when Chickie confronts him about his drinking problem, and Russell is caught in the middle when he attempts to defuse an ongoing domestic battle between a neighbor and his wife.

Lydia and her new partner, Detective Rene Cordero, investigate the disappearance of an elderly man. Meanwhile, Nate and Sammy investigate a freeway shooting and inadvertently threaten the integrity of a major undercover operation. Elsewhere, Chickie finds herself in the middle of a mini-riot after her new partner panics and takes a shot at a 13-year-old boy.

John and Ben are the first responders to a gruesome triple homicide in Hancock Park, where a mother is bludgeoned to death and her young daughters are found bound, gagged, raped and dead in their bedrooms. In investigating the case, Lydia and Rene realize they have two very different investigation styles. Meanwhile, Sammy continues to have problems with his wife and Ben confronts John about his increasing reliance on pain pills.

Ben, assigned to work on his own for the first time, responds to a call about a 9-year old girl who wandered off and finds himself emotionally distraught over a woman who is being harassed by an ex-lover with a tendency towards violence against women. Lydia and Rene spend the day checking assisting Brianna, the mother of the baby found crawling down the street. Over in Gangs, Detective Gil Puente leads the task force to take down ex-banger Trinny Day, but clashes with Sal and Sammy who find him loud and abrasive. When Chickie is assigned to ride with John for the day, he discovers she has no "command presence" and has lost her edge, suggesting it may be time for her to get out of police work.

Lydia and Rene are called to Compton to investigate the murder of a young SULA student, only to discover that the girl was not exactly who her parents claimed she was. Meanwhile, John takes Ben with him to Indian Wells to attend the funeral of his friend, a cop who committed suicide. On the way back, they stop at the rehab center to check in on Dewey.

After two men murder a man in a crowded nightclub, Nate discovers his daughter was an eyewitness. Meanwhile, tensions rise between Sal and Gil on the gang task force, and Sammy confronts his wife, Tammi, after discovering she has been photographing gang members.

John, Ben, and Chickie are sent out as part of a maximum deployment to capture the “Canyon Rapist,” a perp who's been impersonating a cop. Russell is back, but on light duty as he continues to struggle with his recovery after being shot. While working a double-homicide with his old partner, Lydia, he discovers he's no longer up for the job. Sal plans a special night out to commemorate his wife's promotion, but their evening is ruined when their daughter reveals Sal’s affair with TV news reporter Mia Sanchez. Chickie gets a chance to prove to herself -- and others -- that she still has what it takes to be a good cop.

Lydia endures a stressful adjustment period with her new partner, Detective Josie Ochoa, as they investigate the disappearance of an illegal immigrant. Meanwhile, Russell must learn to adapt to a new desk job, Sammy and Nate look for the killer of two gangbangers, Ben catches the eye of an attractive woman, and John struggles with increasing back pain.

Lydia and Josie are assigned to assist Sammy and Nate with a series of gang murders. John discovers he can't stand the "new and improved" Dewey, fresh from rehab, when he's forced to ride with him in order to evaluate whether Dewey is competent to return to duty. Ben must then ride with Chickie, who drives him nuts with her very personal questions about his private life and his new relationship with the "cop groupie" he's been seeing.

Ben has a confrontation with his mother’s rapist after the man is released from prison and is devastated when his mom reveals some information she's been keeping secret for years. Meanwhile, Lydia and Josie continue to bump heads, and Sammy deals with his wife’s betrayal.

Ben learns the hard way, some valuable lessons about the importance of following established procedure and protocol, Lydia's current murder investigation takes an unexpected turn, and Nate and Sammy are unprepared when a group of people they encounter turns into a violent mob.

When a rape victim hires a lawyer and refuses to talk with the two detectives unless her lawyer is present, Lydia and Josie want to know why. Meanwhile, John strikes up an unlikely friendship with a boy who returned home after a sleepover, only to discover his parents had packed up and moved during the night, abandoning him and leaving him to fend for himself, Later on, John begins to feel the stress over the prospect of testifying at his father's upcoming parole board hearing, but he doesn't realize his testifying will open up an emotional can of worms that he's not prepared for.

While working a celebrity murder, Lydia discovers the pros and cons of being in the spotlight. Meanwhile, six weeks after Nate's murder, Sammy finally returns to street patrol. Elsewhere, John voices his anger over the department's waste of money, man-power, and hours spent guarding the home of a celebrity suspected of murdering his estranged wife from the reporters trying to get a scoop.

When crime scene photos from the case of the celebrity suspected of murdering his estranged wife show up on the internet, all of the first responders are suspended, including Lydia and Josie. Further investigation proves the photos were the unauthorized pictures taken by Lydia using her phone, and she's shocked when she learns who was responsible for the leak. Meanwhile, Sammy fails to pick the right guy from the line-up for Nate's murderer, and Dewey and Chickie are involved in a high-speed chase that ends in disaster.

Sammy seeks answers in the desert, Lydia and Josie babysit an important alcoholic witness before a murder trial, and John and Ben go against procedure to help a child in need.

As John's back pain worsens, so does his drug problem, and Ben, about to finish his training period, is gravely concerned about him. Meanwhile, when a man wielding an automatic weapon goes on a killing spree inside a factory, Josie gets injured and Lydia must go after him alone. Elsewhere, Sammy decides to take a more active roll in the upcoming birth of his child..

On Ben's final day as a rookie, he decides to force John to get the surgery he needs to relieve his back pain which, in turn, will end his need for painkillers. Meanwhile, Josie discovers that her son is the man Lydia has been seeing. Elsewhere, Sammy decides to go back to regular patrol.