Kara Thrace seeks her own path in the search for Earth while the Cylons face the threat of sentient Centurions. A member of Galactica's crew learns a dark secret about a loved one.

President Roslin faces challenges as the Colonials plot with Cylon rebels to destroy a Resurrection Hub.

With President Roslin missing along with the cylon base ship a new president must be decided on. Galactica goes on a search for the missing base ship and the Galactica crew members that were on board.

Loyalties are made clear when Gaeta orchestrates a coup d'état of the Adama regime. Both prisoners and lives are taken as many of the fed up people work to take over Galactica.

President Roslin faces off against Tom Zarek and Lt. Gaeta as they try to take control of the Colonial fleet. Both the alliance with the rebel Cylons and Adama's life are at stake.