After Lucy's claims about who Jules is leaves her unsettled, Jules begins to question her past, taking Wes on a journey to investigate. Lucy and Jack chase down a lead that may reveal Jules' origin. And Kira confronts Darros about Lucy.

Kira's research as a graduate student captures the attention of Paul Darros as well as Professor Eleanor Miller, setting in motion a life marked by love, achievement, and struggle that carries enormous ramifications for Lucy many years later.

Kira teams up with Lucy to chase down a lead about Jules, all while Kira navigates problems at home. After being kidnapped and taken to Paul Darros' mysterious compound, Jules attempts to escape captivity with the help of an unusual new friend.

Kira's life starts falling apart when Lucy discovers another printout of Eleanor, and Lucas learns the truth about what happened to his mother. Jules sees another chance to escape from Darros, but he and Xander stay one step ahead of her.

Craig reveals a secret to Lucy; Xander on the memory drug makes him question whether he wants to carry on Darros' legacy; as Kira gets closer to uncovering Darros' master plan, a security breach could cause her to lose everything she's discovered.