Lifeline, a Brussels-based resistance organisation smuggling downed RAF fliers back to Britain, is endangered when Luftwaffe Major Erwin Brandt blows one of their safehouses. The Lifeline leaders, Lisa Colbert and café proprietor Albert Foiret, are suspicious when London sends Flight Lieutenant John Curtis, who had previously gone down the line with Lisa, to help co-ordinate their activities. Brandt, meanwhile, also finds assistance from Gestapo Sturnbannfuhrer Ludwig Kessler, transferred from Berlin to crack down on the evasion lines.

The Jewish population of Brussels is being systemically deported according to colour-coded identification papers developed by the Germans. Curtis becomes involved with a Jewish family hiding out in the city and asks Lisa if Lifeline can help, but Lisa is unwilling to put the line, which is supposed to help RAF evaders not Jews, at risk. Curtis himself is still under suspicion by Albert, when Lisa’s uncle, bank manager Gaston Colbert, identifies Belgian banknotes he brought over from London as forgeries. Kessler obtains some of the notes, and realises that they are too accurate to be real and have been made by the Belgian treasury plates, which were evacuated to London before the German invasion. He believes that the notes will bring him closer to Lifeline.

An American airman has written a magazine article about his experiences being shot down in Europe and getting back home with the help of the evasion lines. British Intelligence has blocked the article’s publication, but Brandt has managed to obtain a copy from German intelligence based in America. Studying the article, he begins to deduce the probable route taken during the evasion and travels to the Pyrenees, where he enlists the aid of Malaud, a French policeman, to close in on the line. Meanwhile, Lisa is also in the area, taking some evaders down the very route Brandt is so close to tracking down.

Flight Lieutenant Peter Romsey, a RAF navigator, is one of Lifeline’s latest customers and London are anxious for his swift return – his technical knowledge of the Oboe navigation system is vitally important. Natalie takes Romsey and other evaders down the line, but Romsey is separated from the others and takes the wrong train, finding himself lost in the French countryside. As Lifeline tries to track him down, Romsey finds unexpected assistance from Hugh and Dorothy Neville, an expatriate English couple. A celebrated author utterly disillusioned with the war, Neville must decide where his true loyalties lie when Romsey turns up on his doorstep.

When Lisa Colbert is arrested whilst meeting a contact near Paris, the whole future of Lifeline is put at risk. This also leads to her uncle being arrested and interrogated by Kessler. Whilst in prison, Lisa must decide whether she can trust her fellow inmates who claim to know who she is and how they can help her escape. These events will lead to tragedy, both personal and for Lifeline.