Led by Majors Cleven and Egan, the 100th Bomb Group arrives in England and joins the 8th Air Force's campaign against Nazi Germany.

The 100th bombs German U-boat pens in Norway. With the help of Lt. Crosby's navigating, a damaged B-17 struggles to get back to Britain.

The group participates in its largest mission to date—the bombing of vital aircraft manufacturing plants deep within Germany.

Lt. Rosenthal joins the 100th just as one of its crews reaches a milestone. The U-boat pens at Bremen become a target for the second time.

Rosie's next mission signals a significant shift in the 100th's bombing strategy. Crosby receives a promotion, but it comes with a high price.