After the Orville crew defeats a Krill attack on a brand-new colony, the Union sends Ed and Gordon on a dangerous undercover mission to infiltrate a Krill ship and obtain a copy of the Krill bible, the "Anhkana," to better understand the enemy.

One of Bortus's old lovers is assigned to help upgrade the ships defences and makes a surprising admission to Talla.

The crew must contend with the disastrous fallout from Kelly's decision.

The Orville discovers a Kaylon with a very special ability.