Julien, Maurice and Pancho regroup aboard a submarine and ponder their next move, while a captive Mort plots to take down the tyrannical new king.

A band of disgruntled Russian space chimps may be Julien's best hope for fighting Koto. Mort finds Timo locked in his pod and begs him for help.

Julien and Karl set out to stop Koto's wedding to the Crocodile Princess. Clover deals with Koto her own way. Mort fights to bring back "Smart Mort."

Nearly vanquished by Koto's troops, Julien rallies Mort's army, the bell snails, the space chimps, Clover and other loyal friends for one last hurrah.