Julien, Maurice and Pancho regroup aboard a submarine and ponder their next move, while a captive Mort plots to take down the tyrannical new king.

As Julien and company head back to Madagascar, their submarine runs into trouble. Mort puts on a play for Koto to cover up the captives' escape.

Clover rethinks her decision to join Pam's cult. Mort uses Timo's radio to contact Julien and company, who are now in trouble with the chimps.

A colony of bell-shaped creatures hails Maurice as their Chosen One. Stuck in limbo, Julien stands trial for his life. Mort battles Morticus Khan.

Nearly vanquished by Koto's troops, Julien rallies Mort's army, the bell snails, the space chimps, Clover and other loyal friends for one last hurrah.