Small-town reverend Ezekiel Brown enlists outlaw Benny the Teen to lead his people west along the Oregon Trail. Repressed prairie wife Prudence Aberdeen responds to the freedom of the trail, pulling her away from her wealthy husband Todd.

Ezekiel signs up for a buffalo hunting expedition to prove his manhood. Meanwhile, Benny teaches Prudence how to be a bandit.

The wagon train stops in the sinful town of Branchwater, luring Ezekiel into a night of debauchery and temptation. Meanwhile, Benny finds his favourite saloon has transformed into an inauthentic tourist trap.